 Research
 Open Access
Stand management optimization – the role of simplifications
 Timo Pukkala^{1}Email author,
 Erkki Lähde^{2} and
 Olavi Laiho^{2}
 Received: 31 May 2013
 Accepted: 4 October 2013
 Published: 26 February 2014
Abstract
Background
Studies on optimal stand management often make simplifications or restrict the choice of treatments. Examples of simplifications are neglecting natural regeneration that appears on a plantation site, omitting advance regeneration in simulations, or restricting thinning treatments to low thinning (thinning from below).
Methods
This study analyzed the impacts of simplifications on the optimization results for Fennoscandian boreal forests. Management of pine and spruce plantations was optimized by gradually reducing the number of simplifying assumptions.
Results
Forced low thinning, cleaning the plantation from the natural regeneration of mixed species and ignoring advance regeneration all had a major impact on optimization results. High thinning (thinning from above) resulted in higher NPV and longer rotation length than thinning from below. It was profitable to leave a mixed stand in the tending treatment of young plantation. When advance regeneration was taken into account, it was profitable to increase the number of thinnings and postpone final felling. In the optimal management, both pine and spruce plantation was gradually converted into unevenaged mixture of spruce and birch.
Conclusions
The results suggest that, with the current management costs and timber price level, it may be profitable to switch to continuous cover management on medium growing sites of Fennoscandian boreal forests.
Keywords
 Plantation management
 Continuous cover forestry
 Evenaged management
 Mixed stands
Background
Most silvicultural instructions have been developed for onespecies evenaged stands (e.g., Anonym 2006). In the best case these instructions are based on simulations and optimizations. If optimization is used the instructions should show the best possible management for a certain objective function. However, caution may be necessary when applying the developed instructions in forestry practice. This is because the models used in calculations may not accommodate all relevant elements of stand dynamics, or the solved optimization problems may simplify reality. The growth and yield models or the optimization problems may be restricted to evenaged stands, unevenaged stands, onespecies stands, low thinnings, etc. It is not uncommon that some important elements of stand dynamics like advance regeneration are ignored when stand development is simulated (e.g., Valsta 1992; Pukkala and Miina 1997).
Several growth and yield simulators are driven by a model that predicts dominant height as a function of stand age and site index (e.g., Shater et al. 2011; Guzmán et al. 2012). Use of the dominant height model assumes that dominant height is not reduced in thinning. Therefore, high thinnings are ruled out also from simulations and optimizations (e.g., Palahí and Pukkala 2003; PasalodosTato et al. 2009). Because of this, it is not known whether the solutions of the optimization problems really indicate the optimal stand management. Several studies suggest that high thinnings may be better than low thinnings when economic profitability is maximized (e.g., Haight et al. 1985; Haight and Monserud 1990; Hyytiäinen et al. 2005).
Most optimizations have been done for pure onespecies stands (e.g., Hyytiäinen et al. 2004; 2005; Tahvonen 2011). However, many stands and plantation sites get natural regeneration of several species (Miina and Saksa 20062008). There may also be advance regeneration from the previous rotation which remains alive after clearfelling. Therefore, it is often possible to obtain a mixed stand even when only one species is planted. The possibility to utilize naturally regenerated admixtures of secondary species is seldom taken into account in optimization. Since the amount of natural regeneration is hard to predict it is often assumed that it does not exist, or it is supposed that seedlings other than the planted ones are removed in the tending treatments of the young plantation. Sometimes the growth and yield model, or the problem formulation used in optimization, has been developed for onespecies stands, which means that admixtures of other species must be ignored because of “technical” reasons. Even if the model accommodates several species it may not describe the interactions between species in a proper way.
Advance regeneration is seldom predicted in the optimizations for evenaged stands. However, if regeneration and ingrowth are plentiful, they may have a major impact on the optimal stand management. It might be more profitable to release the understorey trees instead of clearcutting all trees and establishing a new stand by planting. The reason for ignoring ingrowth may be the lack of recruitment models or a belief that it is an insignificant element of stand dynamics. However, understory reinitiation is a well recognized stage of the natural dynamics of boreal forests (Oliver and Larson 1996). Sooner or later, natural regeneration begins to appear in all evenaged boreal stands. Both the start and the speed of regeneration depend on overstorey management. If the overstorey consists of shadeintolerant pioneer species, such as pine (Pinus sylvestris), aspen (Populus tremula) and birch (Betula pendula and B. pubescens), more shadetolerant species like spruce (Picea abies) typically appear in the stand as understorey. This kind of gradual species succession often begins at quite a young age.
This study analyzed the consequences of the simplifying assumptions listed above in the management of boreal forests. The aim was to find out how the picture about optimal stand management changes when the restrictions and simplifications are gradually removed. Do numerically derived proposals for optimal management change when more realism is added to optimization and to the simulation of stand dynamics? This question was analyzed in spruce and pine plantations established on medium site in Central Finland.
Methods
Existing models for recruitment, tree growth and survival were used to simulate stand dynamics in alternative management schedules. The amount of established regeneration in a plantation site was predicted with the models of Miina and Saksa (20062008). Their models predict the number of surviving individuals of the plantation species, as well as the amounts of natural regeneration of pine, spruce, birch, and hardwoods other than birch. The prediction depends on site characteristics, site preparation method and the used regeneration method (planting, sowing, or natural regeneration from seed or shelter trees).
In addition to the number of seedlings, the models of Miina and Saksa also predict the mean height of the seedlings three years after planting. Since the residual standard deviations of the height models are also reported, it is possible to generate initial size variation among seedlings. In this study, 15 seedlings of different sizes (15 size classes) were generated to represent the planted species (spruce or pine). In addition, 5 seedlings were generated to represent the other conifer (pine or spruce), 5 seedlings for silver birch (B. pendula), 5 seedlings for pubescent birch (B. pubescent), and 5 seedlings for hardwoods other than birch. As a result, a 3yearold seedling stand corresponding to an average conifer plantation on medium site was obtained.
A tending treatment was simulated at the age of 7 years. It was first assumed that species other than the planted one are completely removed. Then, other optimizations were conducted in which a mixture of pine, spruce and birch was left to continue growing. Hardwoods other than birch were removed completely. Trees were left in all size classes of the plantation species but smaller classes were thinned more than larger ones. Other species were treated with uniform thinning. As a result, the tending treatment reduced the size variation of seedlings only slightly. The tending of the spruce plantation left 330 pines, 1028 spruces, 330 silver birches and 330 pubescent birches per hectare. In the pine plantation, 1525 pines, 300 spruces, 90 silver birches and 90 pubescent birches were kept. The number of birches was lower because of the adverse effect of a dense birch cover on pine development. This is because birches easily overtop pines, which reduces the productivity of the stand. This was also noted in preliminary simulations.
The further development of trees was predicted with the models of Pukkala et al. (2013). Their model set consists of individualtree diameter increment model, individualtree survival model, and models for ingrowth. The ingrowth models are based on stands in which the basalareaweighted mean diameter is at least 10 cm. Accordingly, in the simulations of this study, ingrowth was predicted once the mean tree diameter exceeded 10 cm. Species interactions are included in all models. The diameter increment models predict, among other things, that pine and birch competitors reduce the growth of spruce less than spruce competitors. The ingrowth models predict spruce ingrowth also in pure pine and birch stands, which corresponds to the natural dynamics of boreal forests.
The assortment volumes of removed trees were calculated using the taper models of Laasasenaho (1982). Tree height was required in volume calculations and it was predicted with the models of Pukkala et al. (2009). The top diameters of timber assortments were as follows:

Pine: 15 cm for saw log, 8 cm for pulpwood

Spruce: 16 cm for saw log, 8 cm for pulpwood

Birch: 16 cm for saw log, 9 cm for pulpwood
The crosscutting of each removed tree was simulated, taking into account the minimum piece lengths of different timber assortments. A certain percentage of saw log volume was moved to the pulpwood component to mimic the effect of quality defects. The deduction in saw log volume was 10% for pine, 5% for spruce, 15% for silver birch and 20% for pubescent birch.
Roadside timber prices were used to calculate the income from harvests. The roadside prices were 60 €/m^{3} for pine and spruce saw log, 50 €/m^{3} for birch saw log, and 30 €/m^{3} for pulpwood. To calculate net income, harvesting costs were subtracted from the roadside value of harvested trees. The models of Valsta (1992) were used. According to these models the harvesting cost per removed cubic meter decreases with increasing mean size of harvested trees and increasing volume (m^{3}/ha) of the harvest. Thinnings have a higher harvesting cost per cubic meter than clearfelling if the removed volume and mean size of removed trees are the same. Stand establishment cost in year 0 was 1400 €/ha. The tending cost in year 7 depended on the number and diameter of the removed seedlings, but it was close to 270 €/ha in all cases.
A set of optimizations was done for spruce and pine plantations growing on medium site (mesic site, Myrtillus type). This site type is by far the most common fertility class in the southern part of Finland, covering almost 50% of productive forest land. All the main tree species of Finland grow well in this site, making it possible to manage the forest in many different ways.
 1.
All species except the planted one were removed in the tending treatment of the young stand; two thinnings were conducted during the rotation; at most 40% of stand basal area could be removed in thinning; thinnings were conducted as low thinning.
 2.
Otherwise the same but thinning type was not restricted to low thinning.
 3.
Otherwise similar to alternative 2 except that thinning intensity was not restricted.
 4.
Otherwise similar to alternative 3 except that a mixture of pine, spruce, silver birch and pubescent birch was left in the tending treatment of the young stand (at year 7).
 5.
Otherwise similar to alternative 4 except that ingrowth was simulated and the number of thinnings was increased from two to three.
 6.
Otherwise similar to alternative 5 except that the number of thinnings was increased from three to four.
 7.
Otherwise similar to alternative 6 except that the number of thinnings was increased from four to five.
Two thinnings were used in alternatives 1–4 since it has been found that additional thinnings no longer increase the net present value when stand management is optimized without simulating ingrowth (e.g., Pukkala 2006). Alternative 1 corresponds to optimizing the type silviculture that was recommended in Finland for several decades (Anonym 2006). High thinning was forbidden and thinnings could not be too heavy. In the optimization, low thinning was forced by penalizing the solution if the mean tree diameter increased less than 5% as the result of thinning.
Alternative 2 reflects the current situation in which high thinning is permitted but there are regulations that prevent very heavy thinnings. Alternative 3 is a step forward to greater freedom and flexibility, and it may lead to solutions that violate the current forest law of Finland. Henceforth, alternative 1 is referred to as the “lowthinning” alternative and the other alternatives are called as “freethinning” alternatives.
Alternative 4 recognizes the fact that developing mixed stands is often possible even if only one species was planted. Alternatives 5–7 reflect the recent acceptance and increased use of continuous cover management. They allow selective felling and developing the new tree generation from advance regeneration.
The type of thinning was included in the optimization problems by specifying the harvest percentage separately for three different diameter classes. The classes were 10–14 cm, 14–20 cm and > 20 cm for conifers and 10–16 cm, 16–22 cm and > 22 cm for birch. The classification was based on calculations of the absolute and relative value increments of different tree sizes. Trees less than 10 cm in dbh have very high relative value increment during the coming 5–15 years. On the other hand, the road side value of these trees is small and their harvesting is costly. Therefore, it was assumed that trees smaller than 10 cm should never be removed in thinnings. Another diameter class for which the value increment is good is 14–20 cm for conifers and 16–22 cm for birches. This is because trees of this size are approaching the saw log size and their value will therefore increase rapidly. The remaining two diameter classes, 10–14 am (10–16 cm in birch) and > 20 cm (> 22 cm), have much lower relative value increments, which means that removing trees from these classes does not reduce the value increment of the stand as much as cutting trees that are approaching a value threshold.
Based of this rationale, a thinning treatment was specified with three harvest percentages that were optimized separately for pine, spruce, silver birch and pubescent birch. The number of years since planting (first thinning) or previous cutting (other thinnings) defined the timing of the thinning treatment. Therefore, the number of optimized decision variables was 13 (4 × 3 + 1) for each thinning treatment and 1 for the final felling (number of years since the last thinning). A management schedule with 5 thinnings included 5 × 13 + 1 = 66 optimized variables.
The used optimization method was the direct search algorithm proposed by Hooke and Jeeves (1961). In each problem, the direct search was repeated 10 times and the best solution was selected. Every search was started from the best of 100–700 random combinations on decision variables. The number of random searches was increased with increasing number of optimized variables. NPV to infinity (i.e., soil expectation value) with 3% discount rate was maximized in all problems. When calculating the NPV, it was assumed that rotations similar to the simulated one are repeated to infinity.
Results and discussion
Spruce
Net present value calculated with 3% discount rate and optimal rotation length in the optimizations conducted for spruce plantation
Optimization problem  NPV, €/ha  Rotation length, yrs  

10% damage  40% damage  10% damage  40% damage  
1 Low thinning max 40%  408  73  
2 Free thinning max 40%  651  92  
3 Free thinning no limit  845  101  
4 Mixed stand  1088  88  
5 Ingrowth, 3 thinnings  1137  1107  126  118 
6 Ingrowth, 4 thinnings  1187  1116  128  139 
7 Ingrowth, 5 thinnings  1228  1153  131  171 
Pine
Net present value calculated with 3% discount rate and optimal rotation length in the optimizations conducted for pine plantation
Optimization problem  NPV, €/ha  Rotation length, yrs  

10% damage  40% damage  10% damage  40% damage  
1 Low thinning max 40%  847  64  
2 Free thinning max 40%  1190  76  
3 Free thinning no limit  1415  80  
4 Mixed stand  1539  78  
5 Ingrowth, 3 thinnings  1598  1597  112  113 
6 Ingrowth, 4 thinnings  1659  1640  126  130 
7 Ingrowth, 5 thinnings  1663  1649  137  125 
Conclusions
The study presented the first optimizations for pine, spruce and birch mixtures in Finland. Most previous optimizations are for pure onespecies stands (Valsta 1992; Hyytiäinen et al. 2004; Tahvonen 2011), and there are some studies for pinespruce mixtures (Pukkala and Miina 1997; Pukkala et al. 1998; Vettenranta and Miina 1999). The latter studies suggest that pine and spruce mixtures are more productive than pure conifer stands. Optimizations for mixed stands also suggest that mixtures may be healthier than monocultures (Möykkynen and Pukkala 2010). They are also better than monocultures in terms of diversity and scenic value (Silvennoinen et al. 2001). Although yield was not maximized in the current study, the optimizations suggest that mixed stands may be more productive than pure spruce stands. When ingrowth was not included in simulation, the mixed spruce plantation produced 8.7 m^{3}ha^{1}a^{1} whereas the most productive pure stand (free thinning at most 40% of basal area) produced 7.8 m^{3}ha^{1}a^{1}. In the pine plantation, mixed stands were not more productive than a pure pine stand.
This study is also the first one in which advance regeneration was taken into account in the optimization of evenaged stand management. Considering advance regeneration in the analysis has a major impact on the results. It was shown that simplifying assumptions in simulation and optimization may lead to biased conclusions about optimal forest management. It seems that the optimal management of a conifer plantation involves using repeated high thinnings and relying on advance regeneration instead of clearfelling and planting. The maximum number of thinnings tested in this study was five, but most probably the NPV would continue increasing with increasing number of thinnings, implying that it would be optimal to switch to continuous cover management. However, the effect of additional thinnings on NPV would be very small since net incomes discounted from distant future would have only a negligible effect on the total NPV.
The optimizations of this study showed that the first commercial thinning of an evenaged stand should be conducted later than recommended in the thinning guidelines (Anonym 2006), at a basal area of 35–40 m^{2}ha^{1}. The current recommendation is to thin at 25–28 m^{2}ha^{1}. The result agrees with previous research which suggests that the first thinning should be conducted when the stand density is near the self thinning limit (Pukkala and Miina 1998; Vettenranta and Miina 1999; Hyytiäinen et al. 20042005). Earlier thinning, in which only pulpwood is removed, results in a low net return.
The result that high thinning is more profitable than low thinning agrees with several previous studies (Roise 1986; Solberg and Haight 1991; Valsta 1992; Pukkala and Miina 1998; Hyytiäinen et al. 2005). In this study, unrestricted free thinning resulted in 67% (pine) to 166% (spruce) increase in NPV as compared to the optimal lowthinning alternative. Hyytiäinen and Tahvonen (20012003) found that the economic loss to forest landowner may be substantial if official silvicultural instructions are followed instead of the optimal management. The relative differences would most probably be smaller if lower discount rate was used.
Also this study simplified the simulation of stand development and the optimization problem in a few ways. One simplification was that growth variation around model prediction was not simulated, although it would have an impact on the simulation results (Miina 1993; Pukkala et al. 2013). The main influence would be faster differentiation and wider variation of tree size. The consequences of this simplification were reduced by generating size variation in the initial stand. Initial size variation led to the differentiation of tree size, even though residual variation was not simulated. However, differentiation would have been faster if residual variation had been simulated.
It was assumed that ingrowth rate was equal to the model prediction although in reality there is much temporal and spatial variation in regeneration and ingrowth. Simulating regeneration in a stochastic way would correspond better to what happens in nature. However, this would have lead to stochastic optimization, which will be the topic of future studies. The results of this study apply to sites in which the ingrowth is fairly constant and near the average or normal level. Optimal management would be different for stands where ingrowth deviates from this level. However, the small difference between optimization results for 10% and 40% undestorey damage rates suggests that the results are not highly sensitive to the level of ingrowth. Moreover, understorey trees revive at different rates and their growth rate varies much. As a result, temporally irregular regeneration may provide a fairly constant supply of understorey trees to upper canopy layers.
The models that were used in growth simulation (Pukkala et al. 2013) did not describe the thinning effect explicitly; postthinning growth was predicted using the postthinning values of tree and stand characteristics. Previous research suggests that released spruce understoreys may need 4 to 5 years to fully adapt to the new conditions (Metslaid et al. 2005). Therefore, there is a possibility that the postthinning growth was overestimated after heavy high thinnings. The significance of this possibility was analyzed by conducting simulations in which the growth of a highthinning spruce stand was reduced during 5 postthinning years by decreasing the random stand effect of the diameter increment model (by two standard deviations). Twentyyear volume growth was reduced by 11% but 30year growth was only 4% smaller as compared to simulation without growth reduction. Slower initial growth means that competition increases slower resulting in faster growth during later 5years periods.
About 22% of observations behind the used diameter increment model came from a silvicultural experiment in which heavy high thinning was the most common treatment. In addition, a majority of the growth observations of this experiment represented immediate postthinning growth. In the other modeling data sources, the time since previous thinning varied in the same way as it varies in managed Finnish forests (0–40 years). Taking into account that spruce understoreys are usually vigorous under all canopies except a dense and pure spruce canopy, it may be concluded that the impact of thinning stress on the results of this study would remain rather small.
The optimization problem was formulated so that thinning intensity was specified for three diameter classes per tree species and thinning event. Several previous studies have also used three thinning intensities to specify and optimize the type of thinning (Valsta 1992; Pukkala and Miina 1997; Vettenranta and Miina 1999). Using a higher number of diameter classes may have improved the solutions slightly (Hyytiäinen et al. 2005) but, on the other hand, the optimization problem would have become more complicated.
All models and optimization problems simplify reality. Realism should be added as long as it has a significant influence on the results. Those steps that were taken in the current study were all shown to be significant, having a strong impact on our comprehension about the optimal management of boreal forests. Future studies will show whether further steps towards realism are necessary. One improvement, which would most probably have at least some effect, would be to integrate the risks associated to tree growth and survival, and possible changes in timber markets, in the optimization of stand management (Pukkala and Kellomäki 2012). Most probably these analyses would enhance the positive effects that mixed stands and unevenaged stand structures have on the profitability of forestry (Rollin et al. 2005; Pukkala and Kellomäki 2012) since both are ways to share risks (among species, or among cutting events). It would also be interesting to analyze the influence of market trends on optimization results. The price of spruce pulp wood has shown a decreasing trend whereas the price of smallsized birch has increased (Anonym 2012). If the same trend continues, it would be profitable to increase the share of birch more than suggested in this study.
Declarations
Authors’ Affiliations
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This article is published under license to BioMed Central Ltd. This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/2.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited.